Hybrid Tomato Variety &#39;H1991&#39;

ABSTRACT

Hybrid tomato variety ‘H1991’ is described. The tomato variety is a ground-culture hybrid processing tomato variety suitable for machine harvest and is adaptable to the arid climactic conditions of regions such as California, USA, Spain, and Portugal. It is a main season variety with resistance to the three known races of fusarium wilt plus tomato spotted wilt virus. Its thick juice viscosity and medium solids makes it suitable for concentration and its smooth fruit shape lends itself to whole peel applications.

FIELD

This disclosure relates to the field of plant breeding. In particular,this disclosure relates to a new variety of tomato, Solanum lycopersicumvariety denominated ‘H1991’.

BACKGROUND

Breeding improved tomato varieties involves providing genetics that givean advantage to the grower, processor, consumer, or other members of thesupply chain. The improvement may be in the form of field performance,disease resistance, factory performance, or a fruit qualitycharacteristic. For a tomato variety to be suitable to be grown forprocessing, the variety must have a concentrated fruit setting andmaturity, firm fruit, and sufficient rot tolerance to allow early fruitto remain rot-free while later fruit continues to develop and ripen.

Most tomato varieties for commercial processing are hybrids resultingfrom a cross pollination of two true-breeding, inbred parents. Throughthe use of true-breeding lines, a hybrid is produced that often displayscharacteristics of each parent, and often demonstrates characteristicsthat are superior to either parent alone, or that allow a hybrid to maskinadequacies of the individual parents.

Processing tomato varieties combining resistance to verticillium wiltrace 1 (Verticillium dahlia), fusarium wilt races 1 and 2 (Fusariumoxysporum pv lycopersici), bacterial speck race 0 (Pseudomonas syringaepv. tomato), and root knot nematode (Meloidogyne incognita) are highlydesirable in most climates around the world. Moreover, in regions suchas California, the industry has seen a considerable increase in pressurefrom tomato spotted wilt virus (TSWV) and fusarium wilt race 3. Thus,varieties with both resistances are in very high demand by both growersand processors to ensure a productive crop cycle and desirable factoryperformance.

Processing tomato quality parameters differ from those of fruit used inthe fresh market. The processing characteristics are typicallydetermined using a sample of hot-break tomato pulp or juice produced ina consistent manner to those familiar with the art. For example, a fixedmass of tomatoes may be cooked in a microwave oven for several minutesto halt any enzymatic breakdown of the sample, lost water is replaced,and the sample is pulped to remove skins and seed to produce a uniformjuice sample. The juice sample can be analyzed for various qualityparameters important to processing tomato including but not limited togross viscosity measures such as juice Bostwick, soluble solids measuresusing a refractometer (°Brix), measures of acidity and pH, and measuresof color (e.g., a Hunter a/b score). The value of these traits dependson the product that is being commercially produced by the processingfactory. In some instances, a factory will put a higher value on a thickviscosity variety, whereas in other instances, a thin viscosity willmake a superior product and is preferred.

SUMMARY

Provided herein is a new and distinct tomato variety named ‘H1991’ thatproduces very high yields of smooth, medium fruit and demonstrates anadaptability to arid growing conditions around the world. The variety‘H1991’ has resistance to verticillium wilt race 1, three known races offusarium wilt, root knot nematode, bacterial speck race 0, and tomatospotted wilt virus. The fruit of ‘H1991’ have very good internal colorand an average fruit weight of 68 grams. ‘H1991’ presents an advantageover industry standard varieties such as ‘H5608’ in that it carriesresistance to fusarium wilt race 3. ‘H1991’ is designed to allow foronce-over machine harvesting applications during the parts of the seasonwhere extended field holding is not required.

In one aspect, the present disclosure is directed to tomato seeddesignated as ‘H1991’ having ATCC Accession Number PTA-126518. In oneaspect, the present disclosure is directed to a tomato plant, as well asany plant part or portion isolated therefrom, produced by growing‘H1991’ tomato seed. In another aspect, the present disclosure isdirected to a tomato plant or part isolated therefrom having all thephysiological, morphological, and/or genetic characteristics of a tomatoplant produced by growing ‘H1991’ tomato seed having ATCC AccessionNumber PTA-126518. In yet another aspect, the present disclosure isdirected to a tomato plant having all of the physiological andmorphological characteristics of tomato variety ‘H1991’, andspecifically including those listed in Table 1 below, whereinrepresentative seed is deposited under ATCC Accession Number PTA-126518.

In still another aspect, the present disclosure is directed to tomatoseed having at least a first set of the chromosomes of tomato variety‘H1991’, wherein representative seed is deposited under ATCC AccessionNumber PTA-126518. In another aspect, the present disclosure is directedto an F₁ hybrid tomato seed, methods of making F₁ hybrid tomato seed,plants grown from the seed, leaf, ovule, pollen, rootstock, scion,fruit, cotyledon, meristem, anther, root, root tip, pistil, flower,stem, calli, stalk, hypocotyla, pericarp, or portion thereof isolatedtherefrom having ‘H1991’ as a parent, wherein ‘H1991’ is grown from‘H1991’ tomato seed having ATCC Accession Number PTA-126518. Thedisclosure is also directed to a method of producing a tomato plantderived from tomato variety ‘H1991’, including crossing a plant oftomato variety ‘H1991’ with another tomato plant. The method may furthercomprise harvesting seed from the F₁ hybrid tomato seed and/or crossingthe F₁ hybrid tomato plant with itself or another plant to produce seedfrom a progeny plant.

Tomato plant parts include leaf, ovule, pollen, rootstock, scion, fruit,cotyledon, meristem, anther, root, root tip, pistil, flower, stem,calli, stalk, hypocotyl, pericarp, the like, and any portion thereof. Inanother embodiment, the present disclosure is further directed to tomatofruit, stem, leaf, root, root tip, pollen, rootstock, scion, ovule,seed, and flower, and any portion thereof, isolated from ‘H1991’ tomatoplants. In one aspect, the plant part comprises at least one cell fromtomato variety ‘H1991’. In another aspect, the present disclosure isfurther directed to tissue culture of regenerable cells derived from‘H1991’ tomato plants. In one aspect, the tissue culture of regenerablecells has all the physiological and morphological characteristics oftomato variety ‘H1991’. The disclosure is further directed to a tomatoplant regenerated from tissue culture. In another aspect, the disclosureis directed to a protoplast produced from tissue culture and a plantregenerated from the protoplast. At least in some approaches, the plantregenerated from the tissue culture or protoplast has all of thephysiological and morphological characteristics of tomato variety‘H1991’, and specifically including those listed in Table 1.

In another aspect, the disclosure is directed to a method for producinga plant part, which at least in one aspect is tomato fruit, andharvesting the plant part. In another aspect, the plant part comprisesat least one cell from tomato variety ‘H1991’.

The disclosure also is directed to vegetatively propagating a plant oftomato variety ‘H1991’ by obtaining a part of the plant and regeneratinga plant from the plant part. At least in some approaches, theregenerated plant has all of the physiological and morphologicalcharacteristics of tomato variety ‘H1991’, and specifically includingthose listed in Table 1.

In yet another aspect, the present disclosure is further directed to amethod of selecting tomato plants by a) growing ‘H1991’ tomato plantswherein the ‘H1991’ plants are grown from tomato seed having ATCCAccession Number PTA-126518; and b) selecting a plant from step a). Inanother aspect, the present disclosure is further directed to tomatoplants, plant parts, and seeds produced by the tomato plants, where thetomato plants are isolated by the selection method.

In another aspect, the present disclosure is further directed to amethod of breeding tomato plants by crossing a tomato plant with a plantgrown from ‘H1991’ tomato seed having ATCC Accession Number PTA-126518.In another aspect, the tomato plant of tomato variety ‘H1991’ isself-pollinated. In still another aspect, the present disclosure isfurther directed to tomato plants, tomato parts from the tomato plants,and seeds produced therefrom where the tomato plant is isolated by thebreeding method.

In another aspect, the disclosure relates to a plant of tomato variety‘H1991’ comprising a transgene and/or a single locus conversion, and anyseeds or plant parts isolated therefrom. The disclosure also relates tomethods for preparing a plant of tomato variety ‘H1991’ comprising atransgene and/or a single locus conversion. In one aspect, the plantcomprising a single locus conversion and/or transgene has all thephysiological and morphological characteristics of tomato variety‘H1991’.

BRIEF DESCRIPTION OF THE DRAWINGS

The patent or application file contains at least one drawing executed incolor. Copies of this patent or patent application publication withcolor drawings will be provided by the office upon request and paymentof the necessary fee.

FIG. 1 illustrates fruit of tomato variety ‘H1991’.

FIG. 2 illustrates fruit and general plant structure of tomato variety‘H1991’.

DETAILED DESCRIPTION

Described herein is the new and distinct tomato variety named ‘H1991’that was developed to provide a ground-culture hybrid tomato variety(i.e., not grown on stakes) that is suitable for machine harvest, andare adaptable to the climactic conditions of regions such as California,USA, Portugal, and Spain.

Processing tomato varieties combining resistance to verticillium wiltrace 1 (Verticillium dahlia), fusarium wilt race 1 and 2 (Fusariumoxysporum pv. lycopersici), bacterial speck, root knot nematode(Meloidogyne incognita) are highly desirable in most climates around theworld. Varieties with resistance to fusarium wilt race 3 and tomatospotted wilt virus are highly desirable in California as the incidenceof these diseases throughout the state continues to increase.

Tomato plants of ‘H1991’ are resistant to verticillium wilt race 1,three known races of fusarium wilt, root knot nematode, bacterial speckrace 0, and tomato spotted wilt virus. Plants are dark green in colorwith an upright, somewhat open habit that can lead to some of theexposed fruit becoming slightly sunburned as compared to tomatovarieties of the same market class. Mature fruit of ‘H1991’ areextremely firm, very smooth, and medium in size (68 grams). Bothinternal and external color are strong.

Stability of Variety ‘H1991’

The variety is uniform and stable within commercially acceptable limits.As is true with other tomato varieties, a small percentage of variantscan occur within commercially acceptable limits for almost anycharacteristic during the course of repeated multiplication. However, novariants were observed during the three years in which the variety wasobserved to be uniform and stable.

TABLE 1 Characterization of tomato variety ‘H1991’ compared to twoindustry standards, ‘H3402’ and ‘H5608’ Character H1991 H3402 H5608Seedling Anthocyanin in hypocotyl Present Present Present Habit of 3-4Wk seedling Normal Normal Normal Mature Plant Height (cm) 90 n/t n/tGrowth Type Determinate Determinate Determinate Form Normal NormalNormal Size of Canopy Large Large Large Habit Sprawling SprawlingSprawling Stem Branching Profuse Profuse Profuse Branching at CotyledonAbsent Absent Absent # nodes below first 4-7 nodes 4-7 nodes 4-7 nodesinflorescence # nodes between early 1-2 nodes 1-2 nodes 1-2 nodesinflorescence (1^(st)-2^(nd), 2^(nd)-3^(rd)) # nodes between later 1-2nodes 1-2 nodes 1-2 nodes inflorescence Pubescence on younger stemsModerate Moderate Sparse Leaf Type Tomato Tomato Tomato MorphologyCompound Compound Compound with major with major with major and minorand minor and minor leaflets leaflets leaflets (image 2) (image 2)(image 2) Margins of Major Leaflets Shallowly Shallowly ShallowlyToothed Toothed Toothed Marginal Rolling or Wiltiness Slight ModerateModerate Onset of Leaflet Rolling Midseason Midseason Midseason Surfaceof Major Leaflets Rugose Rugose Rugose Pubescence Normal Normal NormalInflorescence Type Forked Forked Forked # flowers in inflorescence 5 5 6average Leafy or “running” Absent Absent Absent inflorescence FlowerCalyx Normal Normal Normal Calyx-Lobes Shorter than Shorter than Shorterthan corolla corolla corolla Corolla Color Old Gold Yellow Yellow Stylepubescence Sparse Sparse Sparse Anthers Fused/Tubed Fused/TubedFused/Tubed Fasciation Absent Absent Absent 1st flower of 2nd or 3rdAbsent Absent Absent Inflorescence Fruit Typical shape Blocky Blockyoval Blocky oval in longitudinal section Shape of transverse sectionRound Round Round Shape of stem end Indented Indented Indented Shape ofblossom end Flattened Flat Flat Shape of pistil scar Dot Dot DotAbscission layer Absent Absent Absent Point of detachment at calyx Atcalyx At calyx fruit at harvest attachment attachment attachment Lengthof pedicel (cm) n/a n/a n/a (joint to calyx attachment) Length of maturefruit 6.25 5.7 5.7 (stem axis) (cm) Diameter of fruit at widest 5.1 4.34.2 point (cm) Weight of Mature Fruit (g) 68 65 77 Number of Locules 2 &3 2-4 2-4 Fruit Surface Smooth Smooth Smooth Fruit Base Color LightMedium Yellow (Mature Green Stage) Green Green green Fruit Pattern(mature green Uniform Uniform Uniform stage) Shoulder color if differentn/a n/a n/a from base Fruit color full ripe Red Red Red Flesh color fullripe Red Red Red Flesh color Uniform Uniform Uniform Locular gel colorof table-ripe Red Red Yellow fruit Ripening Uniform Uniform Uniform StemScar Size Small Small Small Core Coreless Coreless Present EpidermisColor Yellow Yellow Yellow Epidermis Normal Normal Normal EpidermisTexture Average Average Tough Thickness or Pericarp (mm) 6.4 7 5.9Resistance to Fruit Disorder Not tested Not tested Not tested Diseaseand Pest Reactions Viral Diseases Cucumber Mosaic n/t n/t n/t Curly Topn/t n/t n/t Potato-y Virus n/t n/t n/t Blotch Ripening n/t n/t n/tTobacco Mosaic Race 0 n/t n/t n/t Tobacco Mosaic Race 1 n/t n/t n/tTobacco Mosaic Race 2 n/t n/t n/t Cracking, Concentric n/t n/t n/tTobacco Mosaic Race 2² n/t n/t n/t Tomato Spotted Wilt ResistantSusceptible Resistant Tomato Yellows n/t n/t n/t Gold Fleck n/t n/t n/tOthers n/t n/t n/t Bacterial Disease Bacterial Canker Susceptible Min.resistant Susceptible (Clavibacter michiganense) Bacterial Soft Rot n/tn/t n/t (Erwinia carotovora) Bacteria Speck race 0 Resistant ResistantResistant (Pseudomonas tomato) Bacterial Spot Susceptible SusceptibleSusceptible (Xanthomonas spp) Bacterial Wilt n/t Susceptible n/t(Ralstonia solanacearum) Other Bacterial Disease n/t n/t n/t FungalDisease Anthracnose n/t Susceptible n/t (Colletotrichum spp.) Brown RootRot or Corky n/t Susceptible n/t Root (Pyrenochaeta lycopersici) CollarRot or Stem Canker Susceptible Susceptible Susceptible (Alternariasolani) Early Blight Defoliation Susceptible Min. resistant Susceptible(Alternaria solani) Fusarium Wilt Race 1 Resistant Resistant Resistant(F. oxysporum f. lycopersici) Fusarium Wilt Race 2 Resistant ResistantResistant (F. oxysporum f. lycopersici) Fusarium Wilt Race 3 ResistantSusceptible Susceptible (F. oxysporum f. lycopersici) Grey Leaf Spot n/tn/t n/t (Stemphylium spp.) Late Blight, race 0 Susceptible Susceptiblen/t (Phytophthora infestans) Late Blight, race 1 n/t n/t n/t Leaf Moldrace 1 n/t n/t n/t (Cladosporium fulvum) Leaf Mold race 2 n/t n/t n/t(Cladosporium fulvum) Leaf Mold race 3 n/t n/t n/t (Cladosporium fulvum)Leaf Mold Other Races: n/t n/t n/t Nail head Spot n/t n/t n/t(Alternaria tomato) Septoria Leaf spot n/t n/t n/t (S. Lycopersici)Target Leaf spot n/t n/t n/t (Corynespora cassiicola) Verticillium WiltRace 1 Resistant Resistant Resistant (V. dahliae race 1) VerticilliumWilt Race 2 n/t n/t n/t (V. dahliae race 2) Other Fungal Disease n/t n/tn/t Insects and Pests Colorado Potato Beetle n/t n/t n/t (L.decemlineata) Root Knot Nematode (M. sp.) Resistant Resistant ResistantSpider Mites n/t n/t n/t (Tetranychus spp.) Sugar Beet Army Worm n/t n/tn/t (S. exigua) Tobacco Flea Beetle n/t n/t n/t (E. hiritipennis) TomatoHornworm n/t n/t n/t (M. quinquemaculata) Tomato Fruit worm (H. zea) n/tn/t n/t Whitefly (T. vaporariorum) n/t n/t n/t Other n/t n/t n/t

Chemistry and Composition of Full-Ripe Fruits

TABLE 2 Hot-break tomato juice characteristics for new variety ‘H1991’and two check varieties ‘H3402’ and ‘H5608’ ‘H1991’ ‘H3402’ ‘H5608’Serum viscosity (centistokes) 11.4 7.7 10.4 Juice Bostwick (cm) 11.812.6 10.5 Soluble solids (°Brix) 5.3 5.3 5.2 Fruit weight (g) 68 65 77Average of 2 years of trials in California in a total of 21 locations.

TABLE 3 Phenology ‘H1991’ ‘H3402’ ‘H5608’ Fruiting Season Short ShortShort Relative Maturity Medium Medium Medium

TABLE 4 Adaptation ‘H1991’ ‘H3402’ ‘H5608’ Culture Field Field FieldPrinciple use Whole-pack, Whole-pack, Whole-pack, ConcentratedConcentrated Concentrated Machine harvest Yes Yes Yes Regions ofadaptability California Yes-2 Yes-2 Yes-2 Sacramento/upper SJ valleyCalifornia lower SJ Yes-1 Yes-3 Yes-1 valley Northeastern USA No Yes-1No If more than one category applies, they are listed in rank order.

Comparison of ‘H1991’ to Closest Varieties

Data in Table 1 is based primarily upon trials conducted inCollegeville, Calif. from two replications, non-staked, in a researchplot environment. Data in Tables 3 and 4 are based upon observationsmade in trials throughout California and Ontario, Canada. Comparisonsamong varieties for processing traits (Table 2) were done over two yearsof side-by side testing throughout California. Disease resistance andadaptability assessments are based upon DNA markers associated with thedisease resistance when available or numerous observations collected inregions/climates with specific disease pressure, specifically for ripefruit rots, bacterial spot, bacterial canker, early blight, and lateblight, including Ontario, Canada.

Both ‘H1991’ and ‘H5608’ have similar crop maturities, in the mid-seasonclass, yet there are several characteristics that can distinguish‘H1991’ from the commercial variety ‘H5608’. ‘H1991’ has a blockier,fruit shape than ‘H5608’. The fruit color of ‘H1991’ is very good, and adeeper red color than ‘H5608’, particularly in its locular gel.Agronomically, the major advantage of ‘H1996’ is that it carriesresistance to Fusarium oxysporum race 3, which is lacking in ‘H5608’.Varieties with good field performance, thick (low) juice Bostwick, andwhich also carry resistance to fusarium wilt race 3 are currently highlyvalued by the processing tomato industry.

Further Embodiments

Additional methods provided herein include, without limitation, chasingselfs. Chasing selfs involves identifying inbred plants among tomatoplants that have been grown from hybrid tomato seed. Once the seed isplanted, the inbred plants may be identified and selected due to theirdecreased vigor relative to the hybrid plants that grow from the hybridseed. By locating the inbred plants, isolating them from the rest of theplants, and self-pollinating them (i.e., “chasing selfs”), a breeder canobtain an inbred line that is identical to an inbred parent used toproduce the hybrid. Accordingly, another aspect of the disclosurerelates a method for producing an inbred tomato variety by: plantingseed of the tomato variety ‘H1991’; growing plants from the seed;identifying one or more inbred tomato plants; controlling pollination ina manner which preserves homozygosity of the one or more inbred plants;and harvesting resultant seed from the one or more inbred plants. Thestep of identifying the one or more inbred tomato plants may furtherinclude identifying plants with decreased vigor, i.e., plants thatappear less robust than plants of the tomato variety ‘H1991’. Tomatoplants capable of expressing substantially all of the physiological andmorphological characteristics of the parental inbred lines of tomatovariety ‘H1991’ include tomato plants obtained by chasing selfs fromseed of tomato variety ‘H1991’.

One of ordinary skill in the art will recognize that once a breeder hasobtained inbred tomato plants by chasing selfs from seed of tomatovariety ‘H1991’, the breeder can then produce new inbred plants such asby sib-pollinating, or by crossing one of the identified inbred tomatoplant with a plant of the tomato variety ‘H1991’.

The disclosure further includes introducing one or more desired traitsinto the tomato variety ‘H1991’. For example, the desired trait mayinclude male sterility, male fertility, herbicide resistance, insectresistance, disease resistance, and drought resistance.

The desired trait may be found on a single gene or combination of genes.The desired trait may be a genetic locus that is a dominant or recessiveallele. The genetic locus may be a naturally occurring tomato geneintroduced into the genome of a parent of the variety by backcrossing, anatural or induced mutation, or a transgene introduced through genetictransformation techniques. For a genetic locus introduced throughtransformation, the genetic locus may comprise one or more transgenesintegrated at a single chromosomal location. Accordingly, the disclosureprovides tomato plants or parts thereof that have been transformed withone or more transgenes (i.e., a genetic locus comprising a sequenceintroduced into the genome of a tomato plant by transformation) toprovide a desired trait. In one aspect, the one or more transgenes areoperably linked to at least one regulatory element. In one aspect, thetomato plant or plant part comprising a transgene has all thephysiological and morphological characteristics of tomato variety‘H1991’.

The gene(s) may be introduced to tomato variety ‘H1991’ through avariety of well-known techniques, including for example, molecularbiological, other genetic engineering, or plant breeding techniques,such as recurrent selection, backcrossing, pedigree breeding, molecularmarker (Isozyme Electrophoresis, Restriction Fragment LengthPolymorphisms (RFLPs), Randomly Amplified Polymorphic DNAs (RAPDs),Arbitrarily Primed Polymerase Chain Reaction (AP-PCR), DNA AmplificationFingerprinting (DAF), Sequence Characterized Amplified Regions (SCARs),Amplified Fragment Length Polymorphisms (AFLPs), and Simple SequenceRepeats (SSRs) (also referred to as Microsatellites)), enhancedselection, genetic marker enhanced selection, and transformation.Accordingly, tomato seed, plants, and parts thereof produced by suchgenetic engineering or plant breed techniques are also part of thepresent disclosure.

Also provided herein are single locus converted plants and seedsdeveloped by backcrossing wherein essentially all of the morphologicaland physiological characteristics of an inbred are recovered in additionto the characteristics conferred by the single locus transferred intothe inbred via the backcrossing technique. A single locus may compriseone gene, or in the case of transgenic plants, one or more transgenesintegrated into the host genome at a single site (locus). One or morelocus conversion traits may be introduced into a single tomato variety.In one aspect, the tomato plant or tomato plant part comprising a singlelocus conversion has all the physiological and morphologicalcharacteristics of tomato variety ‘H1991’.

DEPOSIT INFORMATION

A deposit of the tomato variety ‘H1991’ is maintained by HeinzSeedCompany, having an address at 6755 CE Dixon St, Stockton, Calif. 95206,United States of America. Access to this deposit will be availableduring the pendency of this application to persons determined by theCommissioner of Patents and Trademarks to be entitled thereto under 37C.F.R. § 1.14 and 35 U.S.C. § 122.

At least 625 seeds of tomato variety ‘H1991’ were deposited on Dec. 2,2019 according to the Budapest Treaty in the American Type CultureCollection (ATCC), P.O. Box 1549, MANASSAS, Va. 20108 USA. The deposithas been assigned ATCC number PTA-126518. Access to this deposit will beavailable during the pendency of this application to persons determinedby the Commissioner of Patents and Trademarks to be entitled theretounder 37 C.F.R. § 1.14 and 35 U.S.C. § 122. Upon allowance of any claimsin this application, all restrictions on the availability to the publicof the variety will be irrevocably removed for the enforceable life ofthe patent.

The deposits will be maintained in the ATCC depository, which is apublic depository, for a period of 30 years, or 5 years after the mostrecent request, or for the effective life of the patent, whichever islonger, and will be replaced if a deposit becomes nonviable during thatperiod.

What is claimed is:
 1. Tomato seed designated as ‘H1991’, representativesample of seed having been deposited under ATCC Accession NumberPTA-126518.
 2. A plant produced by growing the seed of claim
 1. 3. Aplant part from the plant of claim 2, wherein the plant part comprisesat least one cell from tomato variety ‘H1991’.
 4. The plant part ofclaim 3, wherein the part is selected from the group consisting of leaf,ovule, pollen, rootstock, scion, fruit, cotyledon, meristem, anther,root, root tip, pistil, flower, stem, calli, stalk, hypocotyl, pericarp,and portion thereof.
 5. A tomato plant having all the physiological andmorphological characteristics of the tomato plant of claim
 2. 6. A plantpart from the plant of claim 5, wherein the plant part comprises atleast one cell from tomato variety ‘H1991’.
 7. The plant part of claim6, wherein the part is selected from the group consisting of leaf,ovule, pollen, rootstock, scion, fruit, cotyledon, meristem, anther,root, root tip, pistil, flower, stem, calli, stalk, hypocotyl, pericarp,and portion thereof.
 8. A tomato plant having all of the physiologicaland morphological characteristics of tomato variety ‘H1991’, wherein arepresentative sample of seed has been deposited under ATCC AccessionNumber PTA-126518.
 9. A plant part from the plant of claim 8, whereinthe plant part comprises at least one cell from tomato variety ‘H1991’.10. The plant part of claim 9, wherein the part is selected from thegroup consisting of leaf, ovule, pollen, rootstock, scion, fruit,cotyledon, meristem, anther, root, root tip, pistil, flower, stem,calli, stalk, hypocotyl, pericarp, and portion thereof.
 11. Pollen ofthe plant of claim
 2. 12. An ovule of the plant of claim
 2. 13. A tissueculture of regenerable cells from a plant part of claim 3, wherein saidtissue culture of regenerable cells has all the physiological andmorphological characteristics of tomato variety ‘H1991’.
 14. A tomatoplant regenerated from the tissue culture of claim 13, the plant havingall of the physiological and morphological characteristics of tomatovariety ‘H1991’, wherein a representative sample of seed has beendeposited under ATCC Accession Number PTA-126518.
 15. A protoplastproduced from the tissue culture of claim 13, wherein a plantregenerated from the protoplast has all of the physiological andmorphological characteristics of tomato variety ‘H1991’.
 16. A method ofproducing a tomato plant derived from tomato variety ‘H1991’, the methodcomprising crossing the plant of claim 2 with another tomato plant toproduce a F₁ hybrid tomato plant.
 17. The method of claim 16, furthercomprising harvesting seed from the F₁ hybrid tomato plant.
 18. Themethod of claim 16, further comprising crossing the F₁ hybrid tomatoplant with itself or another plant to produce seed from a progeny plant.19. The plant of claim 2, said plant having all the physiological andmorphological characteristics of tomato variety ‘H1991’, and furthercomprising a transgene.
 20. The plant of claim 2, said plant having allthe physiological and morphological characteristics of tomato variety‘H1991’, and further comprising a single locus conversion.
 21. A methodfor producing a tomato fruit, the method comprising: growing the tomatoplant of claim 2 to produce a tomato fruit; and harvesting the tomatofruit.
 22. A method for producing a tomato seed comprising:self-pollinating the tomato plant of claim 2; and harvesting theresultant tomato seed.
 23. A method of vegetatively propagating theplant of claim 2, the method comprising: obtaining a part of the plant;and regenerating a plant from the part, the regenerated plant having allof the physiological and morphological characteristics of tomato variety‘H1991’ listed in Table 1.